Larva: a medium sized plumosus-type; ventral tubules well developed, posterior pair longer. Gular region generally darkened, frontoclypeus dark, but occasionally pale. Pale headed larvae are very difficult to distinuish from C. 'februarius'. Mentum (Figs. 1 - 5) of type II or III, and c2 teeth generally notches on the c1 tooth, i.e. type I, but may be more developed to type IV. Ventromentum with about 33 - 35 striae. Pecten epipharyngis (Fig. 7) with about 12 - 17 teeth. Mandible of type III, with about 12 - 13 striae. Basal segment of antenna (Fig. 8) relatively long and narrow, about 4 - 5 times as long as wide; AR about 2.3 - 3; A2/A1 about 0.18; A4/A3 about 0.9 - 1.2.
Cytology: 4 pairs of chromosomes with the pseudothummi arm combination AE, BF, CD, G. All
chromosomes generally closely paired.
Arm G with an almost terminal nucleolus, with 2 Balbiani
rings; two of the known BRs are not developed, BR1 proximal to the median BR, and BR4 virtually at telomere. No nucleoli in long chromosomes. Low level of polymorphism in arms A, B, C, D, F, and G.
cloA1: 1a - 2c, 10a - 12c, 3i - 2d, 9e - 4a, 13a - 19 as holomelas, etc.
cloA2: approx 1a - 2c, 11a-e, (6?)7 - 9, 2d - 3, 12a-c, (6?)5 - 4, 13 - 19
cloA3: 1a-e, 13e-a, 4 - 9, 2d - 3, 12 - 10, 2c-a, 13f - 19
cloB1: Puff and distal dark bands (groups 7-8) about 1/3 from centromere.
cloB2: Inversion of distal 2/3 of arm, proximal break in or just distal to puff, distal break near end of arm.
cloB3: Small inversion at distal end of the arm. Distal break closer to end than in B2.
cloC1: 1, 14 - 11d, 6b - 2, 15, 8 - 11c, 6gh, 17a - 16, 7d-a, 6fc,Ê17b - 22 (Wülker)
cloC2:
cloD1: 1 - 2, 17 - 12b, 10 - 7, 3g-a, 6 - 4, 12a - 11, 18 - 24, (Kiknadze)
cloD2:
cloE1: 1a - 2b, 5a - 10b, 3e - 2c, 4h - 3f, 10c - 13
cloF1: 1a - 2a, 4a - 10d, 3f - 2b, 11a - 23
cloF2:
Click here for the polytene chromosomes
Pupa: General coloration is dark on the cephalothorax and abdomen. Pupae from Queensland are smaller (abt. 9.37 mm, females; 8.66 mm, males) than those from Victoria (abt. 10.81 mm., females; 9.66 mm., males).
Frontal tubercles are short and conical. Base of respiratory organ 155 x 75 µm. Anterior to this base is a small elevated patch of wartlike tubercles; dorsal to the base is a larger, more elongated patch of smaller tubercles.
Shagreen pattern and chaetootaxy as in Fig. 12 (above).
Spur on eighth segment normally with 3 or 4 spines, range 1 - 6.
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|
|
Females |
|
Males |
|
|
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Mean |
Range |
Mean |
Range |
|
|
Length
(mm) |
9.75 |
7.56
– 10.84 |
9.17 |
7.64 -
10.43 |
|
|
Inner
margin wing case (mm) |
1.98 |
1.44 -
2.22 |
1.92 |
1.68 - 2.09 |
|
|
Cephalic tubercles
(µm) |
|
|
|
|
|
|
Cephalic
bristles (µm) |
39 |
32 - 42 |
43 |
38 - 45 |
|
|
Recurved
hooks on abd. seg. 2 |
95 |
81 - 108 |
74 |
70 - 106 |
|
|
Swim fin
filaments (one side) |
131 |
72 - 173 |
116 |
98 - 154 |
|
Some data on mentum given by Atchley and Martin (1971). Larval morphology and polytene chromosomes figured by Martin (1971a). Sequences of arms A, E and F given by Wülker et al. (1989). Nucleoli and location of C-bands studied by Lentzios & Stocker (1979) and Lentzios et al. (1980).