Larva a medium sized bathophilus-type; length about 11.5 - 15.3 mm (female); ventral tubules well developed, posterior pair longer, length of anterior 0.68 - 1.20 mm; of posterior about 0.56 - 1 45 mm. Gula varying from pale to posterior third darkened (generally paler in more northern populations), frontoclypeus generally pale or very slightly darkened. Larvae are very difficult to distinuish from other members of the oppositus-group. Mentum (Fig. c) with 4th laterals slightly reduced, sometimes to level of fifth laterals (i.e. between type I-II), and c2 teeth well separated, sometimes the c1 tooth is quite long (i.e. type II or IV). Ventromentum (Fig. d) with about 30-38 striae. Pecten epipharyngis (Fig. a) with about 12-18 teeth. Premandible with the normal two teeth, inner tooth up to twice the width of the outer (1.63 - 2.0). Mandible (Fig. e) generally of type II, with about 14-17 striae. Basal segment of antenna (Fig. b) relatively long and narrow, about 3.5-4.6 times as long as wide; A2/A1 about 0.25; A4/A3 about 0.9-1.3.
Cytology: 4 pairs of chromosomes with the pseudothummi arm combination AE, BF, CD, G. All
chromosomes closely paired.
Arm G with a median nucleolus and two nucleoli, the large (probably) BR2 proximal of the nucleolus and the smaller BR3 near the distal end. There appear to be two pieces of BR1, one between the nucleolus and BR2 and the other just distal of the nucleolus. The potential site of BR4 is immediately distal to the BR1 fragment between the nucleolus and BR2. Nucleolus in arm F at about group 19. Polymorphism in arms A, C and D.
psoA1: 1a-e, 6e - 7, 13 - 14c, 3i-f, 9 - 8, 2d - 3e, 1f - 2c, 10 - 11, 6d - 4, 12a-c, 14d - 19 i.e 14c-12c and then 6e-11 from oppA4
psoA2: only as heterozygote
psoB1: Sequence as B2 of oppositus
psoC1: as C. tepperiC1
psoC2: Sex linked and only as heterozygote in some males; generally includes D2
psoD1: 1-2e, 6-8, 19a-c, 14h-16, 2i-g, 18a-g, 12-10d, 3a-d, 9e-b, 2f, 5-3e, 10a-c, 17a-f, 9a, 13-14g, 19d-24 i.e. inv6-2f from tepD1
psoD2: includes C2; sex linked and only as heterozygote in some males
psoE1: 1 - 3e, 10b - 3f, 10c - 13 as oppositus E1
psoF1: 1 - 2a, 10 - 2b, 11 - 23 as oppositus F1
Click here for the polytene chromosomes
Nucleoli and location of C-bands studied by Lentzios & Stocker (1979) and Lentzios et al. (1980) as C. alternans b.
Pupa: Spur with only one or two spines.