Larva: Medium size melanotus-type larva, length about 10.7 (9.7 - 11.3) mm in females; 10.8 (10.2 & 11.3) mm in males. Lateral tubules about 300 - 400 micron. Length of ventral tubules about 1.45 - 1.9 mm (anterior) and 1.9 - 2.5 mm (posterior). Head capsule with slightly darkened gula and frontoclypeus. Mentum (Fig. c) with 4th lateral slightly reduced, almost to the level of the 5th lateral (type I or II), c2 teeth hardly separated from c1 (type I). Ventromentum (Fig. d) with about 35 - 40 striae; distance between inner margins only about 0.2 of mentum width. Basal segment of antenna (Fig. b) about 3.3 to 4.5 times longer than wide; AR about 1.44 - 1.5; A4/A3 about 1.5 - 1.58; ratio of antennal segments 111 : 39 : 11 : 17 : 8 . Pecten epipharyngis (Fig. a) with about 13 - 15 somewhat irregular teeth. Mandible (Fig. e) with third inner tooth darkened and separated (type III), about 19 - 20 striae near base. Anal tubules about 400 - 560 micron in length, tending to be constricted near the middle.
Cytology: 4 pairs of chromosomes with the thummi arm combination AB, CD, EF, G (Only known species of this cytocomplex in Australia). Arm G with a subterminal nucleolus and a BR near the middle of the arm. No nucleolus in the long chromosomes. No polymorphism in the known specimens.
qldA1:
qldB1: Large puff (groups 7) about one third from distal end, with dark bands (group 8) proximal to it.
qldC1: Distinctive band groups 3-4 about one third from distal end.
qldD1:
qldE1: 1 - 3b, 6 - 10b, 3c-a, 5 - 3f, 10c - 13
qldF1:
Click here for the polytene chromosomes.
Pupal exuvia relatively pale, brownish; various characters as in Table below. Spurs generally with only 1 spine, but sometimes with 2.
|
|
|
Females |
|
Males |
|
|
|
Mean |
Range |
Mean |
Range |
|
|
Length
(mm) |
7.36 |
7.07 &
7.64 |
6.97 |
6.67 -
7.32 |
|
|
Inner
margin wing case (mm) |
1.43 |
1.42 &
1.43 |
1.33 |
1.21 -
1.47 |
|
|
Cephalic
tubercles (µm) |
61 |
51 &
72 |
71 |
64 - 82 |
|
|
Cephalic
bristles (µm) |
46 |
43 &
49 |
38 |
37 - 38 |
|
|
Recurved
hooks on abd. seg. 2 |
69 |
64 &
74 |
72 |
64 - 82 |
|
|
Swim fin
filaments (one side) |
78 |
78 &
79 |
62 |
47 - 72 |
|
Larva very similar to that of C. sp. (bathophilus type) but differing in the presence of lateral tubules and a slightly higher AR and ratio of A4/A3.