Larva: a medium sized plumosus-type larva (length about 10.7 - 14.3 mm., lab. reared). Anterior ventral tubules (1.24 - 1.84 mm.) shorter than posterior pair (1.40 - 2.28 mm.). Gula pale or very slightly darkened on posterior third; frontoclypeus slightly dark to dark. Mentum (Fig. b) with square sharp teeth, c2 teeth of central trifid tooth well separated from c1 tooth (type IV), 4th laterals slightly reduced (type I). Pecten epipharyngis (Fig. a) with about 16 - 21 variable but sharp teeth. Ventromentum (Fig. c) with about 29 - 34 striae. Antenna (Fig. d) with a moderately long basal segment, which is about 4 - 4.5 times as long as wide; AR about 1.75 - 1.96. Antennal proportions: 122 : 31 : 10 : 13 : 6. Mandible (Fig. e) with third inner tooth only slightly darkened (Type I), and with about 12 - 14 striae at the base.
Cytology: 4 pairs of chromosomes with the pseudothummi arm combination AE, BF, CD, G. Nucleolus virtually terminal in arm G; well developed BR about one third from the other end, and a smaller BR close to this other end; closely paired. No nucleolus in long chromosomes of Australian specimens, but one near the characteristic bands of arm F in Japanese specimens. Arm A of Australian specimens differs from that of Japanese specimens by a complex inversion, and arm F possibly by a simple inversion.
samA1: 1 - 2c, 10 - 12, 3 - 2d, 9 - 4, 13 - 19 as holomelas(Australia)
samA2: 1a-i, 2k-d, 9e - 4a, 13a - 14i, 3h-i, 12c - 10a, 2c - 1k, 3a-g, 15 - 19 (Japan)
samB1:
samC1:
samD1:
samE1: 1 - 3e, 10b - 3f, 10c - 13 as halophilis, etc.
samF1: 1 -2a, 10d-a, 15 - 11, 2b - 9, 16 - 23 (Japan)
samF2: (possibly) 1 - 2a, 10a-d, 15 - 14e, 9 - 2b, 11 - 14d, 16 - 23 (Australia)
A partial set of polytene chromosomes can be seen here
Pupa: Exuvia length about 7.2 - 7.8 mm. Spur with 1, 2 or sometimes 3 spines.
Chromosome arms A, E and F were described by Wülker et al. (1989) based on Japanese specimens. However, the arm A sequence of Australian specimens is not the same.
Material identified as C. sameonsis from India is also cytologically distinct.