Larva: Known from European material - plumosus-type larva with unpigmented head capsule. c2 teeth of central trifid tooth of mentum relatively well separated (type I-II), as are the 1st laterals.
Cytology: Not known for North American material. European material has 4 pairs of chromosomes with the thummi-cytocomplex combination AB, CD, EF, G.
Arm G relatively short, with a subterminal nucleolus, and a large BR near the other end with a tight constriction in between. Keyl patterns of the bands are difficult to recognize. Polymorphism in arm C in Europe.
Found: California - Dana Meadows, Tuolumne Co. (Grodhaus & Ferrington 1989); Yosemite National Park (Ryser et al. 1985).
Also found in Finland (Type) and Abisko, Sweden, in Europe. The larva, pupa, adult and cytology are given by Ryser et al. (1985). Grodhaus & Ferrington (1989) give notes on the adult of North American specimens.