Species d. C. maturus Johannsen

Adult male
Wing length about 3.64 mm; width about 0.83 mm; VR 1.06.
Foretarsus bearded, BR about 5.
Abdominal pattern similar to that of C. bifurcatus, but the superior volsella is darker and heavier than C. bifurcatus or C. decorus.    Abdominal terga with dark basal fascia.    AR about 3.    Frontal tubercles present, about 25 µm long.    Clypeus with at least 14 setae, scutellum with an anterior row of about 12 long setae, and a main row of about 6 scattered setae.

Pupa:    General color dark with lateral abdominal markings blackish.    Length about 6.30 - 8.88 mm.    Usually with about 1 or 2 spines on the spurs, but some have 3 on one side.    Swim fin withabout 64 - 96 broad flattened setae.

Larva of the plumosus-type, with long ventral tubules, posterior pair usually longer (ant. 0.9 - 3.4 mm, post. 0.9 - 3.7 mm).    Length about 12.5 - 14.8 mm (female), 12.3 - 13.3 mm (male).    Anal tubules about 3 - 4 times as long as wide.    Gular region slightly darkened, frontoclypeus darkened.    Ventral head length 273-304 µm.    Mentum with rather rounded teeth, centre tooth long and narrow with side teeth well separated (type III), 4th laterals barely reduced (type I).    Pecten epipharyngis usually with fairly uniform teeth, occasionally with an interspersed smaller tooth, about 10 - 19 teeth.    Mandible with third internal tooth well developed but not darkened.

Cytology :    4 pairs of chromosomes with maturus arm combination AF, BE, CD, G.
Arm G often partly unpaired, with a subterminal nucleolus and 2 Balbiani rings whose position varies depending on the sequence.    Nucleolus also in arm F.    Polymorphic in all arms except E.
matA1:    1-2c, 10-12, 3-2d, 9-4, 15-13, 16-19
matA2:    1a-e, 2d-3, 12-10, 2c-1f, 9-4, 15-13, 16-19
matA3:    1a-e, 2d-3, 12-10, 16, 13-15, 4-9, 1f-2c, 17-19          i.e. from A2
matB1:    Puff with distal dark bands (groups 7-8) nearer the end of the arm than in whitseli.
matC1:    1a-i, 10-11c, 2-6b, 9f-a, 6hg, 11d-15, 8a-g, 17-16a, 7d-6c, 17b-22
matD1:    1a-i, 15e-11, 3-2, 16-18f, 7d-4, 10-7e, 18g-24
matE1:      1 - 3e, 8 - 5, 9 - 10b, 4 - 3f, 10c - 13;                      i.e. as stigmaterus.
matF1:    1 - 2, 15e - 3, 15f - 23
matF2:    1, 13b - 15e, 2, 13a - 3, 15f - 23

Found: Manitoba - Churchill; Southern Indian Lake (Rosenberg et al., 1984)
            Ontario - Copanspin Farm & Dunrobin, Central Experimental Farm & Hogs Back, Ottawa, Mile 14.3,
            Highway 60, Algonquin Provincial Park.
            Alaska - Potters Marsh, Anchorage Co.
            California - Berkeley, Hayward, Oakland and Strawberry Canyon, Berkeley Hills, Alameda Co.;
            El Cerrito, West Pittsburg, Trout Farm, Concord, Wildcat Creek, nr. El Cerroti, Jewell Lake,
            Tilden Park and Antioch, Contra Costa Co.; state line, Eldorado Co.; Mad River Beach, Humboldt Co.;
            Lakeport and Clear Lake Park, Lake Co.; Susanville, Lassen Co.; Rio Hondo, Montebello,
            and Pasedena, Los Angeles Co.; Lily Lake, Marin Co.; 15 ml nw. Canby, Modoc Co.; 3 ml sw. &
            1 ml. sw. Napa, Napa Co.; Huntington Beach, Orange Co.; Lake Almanor & Lake Davis, Plumas Co.;
            Bergh Ranch, Coachella Valley, Arlington, Good Samaritan Retirement Home, Corona,
            448 Orange St., and UCR Experimental Ponds, Riverside, Palm Springs, Hidden Lake, 3 ml. n. Arlington,
            & Priester Ranch, Norco, Riverside Co.; Ontario & Spring Valley Lake nr Hesperia, Apple Valley,
            San Bernadino Co.; San Luis Obispo & Black Lake Canyon, San Luis Obispo Co.; Redwood City,
            Laurel Creek, marshland pond Millbrae, 19th & Bayshore & 16th Ave., Hayward Park,
            San Mateo, & Atherton, San Mateo Co.; Stanford Uni., Palo Alto & Mountain View, Santa Clara Co.;
            Hat Creek, Fall River Mills, Shasta Co.; Vallejo, Dixon, & Rio Vista, Solano Co.; Cotati, Sonoma Co.
            Indiana - Ridinger Lake, Kosciusko Co.
            Louisiana - Fish Hatchery & Chaplain's Lake, Natchitoches, Natchitoches Ph.
            New Mexico - 2 ml. Bonito Dam, Lincoln Co.
            New York - Ithaca (type locality).
            South Dakota - 3 ml. w. Yankton & Yankton.
            Wisconsin - Trout Lake Limnological Station, Vilas Co.

            Shallow pools, often temporary.

Morphology of larva, pupa and adult described by Sublette & Sublette (1974).    Cytology described by Wülker and Martin (1974), with sequence of arm C1 and arm D1 in Kiknadze et al. (2004).

[ See also C. whitseli | Return to Index | Go to References ]


Modified: 19 July 2009
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Copyright © 2000-2009, Jon Martin.