Species r.  C. 'tigris' Butler and Kiknadze
Now a nomen nudum as the name was published by Martin et al. (2008).

Pupa with about 5 or 6 spines on spurs.

Pupa of C. ‘tigris’
Cephalic tubules (above), spur (below)

Larva a large plumosus-type (length, female: 17.6 - 20.4 mm) with anterior ventral tubule longer (female: Ant. 1.5 - 2.4 mm; post. 1.3 - 2.1 mm).    Gular and frontoclypeus darkened.    Mentum (Fig. c) with a relatively shallow curve, teeth pointed, 4th laterals slightly reduced (type II); c1 tooth broad with short parallel sides, c2 teeth well developed (type II).    Pecten epipharyngis (Fig. a) with about 18 - 19 somewhat irregular teeth.    Ventromentum (Fig. d) with about 35 - 39 striae; anterior margin relatively smooth.    Antenna (Fig. b) with relatively short, broad segment 1, about 2.6 - 2.75 times as long as wide; AR about 2.18; antannal segments 152 : 31 : 10 : 15 : 8 micron.    Mandible (Fig. e) with third inner tooth separated but still relatively pale (type II - III), about 15 striae on inner surface.

Cytology:    2 pairs of long chromosomes, banding pattern often unclear.    Arm combination GAB, FEDC.
Nucleoli in the region of arm G and arm D, i.e. one in each chromosome.    Balbiani rings distal to nucleolus in arm G.    Centromere of arm F, along with adjacent proximal bands of arm E, translocated to distal end of arm E at junction with arm D, where it presumably forms the functional centromere of this arm.

tigA1:     1, 9 - 12, 3 - 2, 8 - 4, 13 - 19
tigB1:     Puff (group 7) with some distal dark bands near end of chromosome
tigC1:     1 - 6c, 11 - 7c, 15 - 12, 7b - 6d, 16 - 22
tigD1:     1 - 3, 11 - 18f, 4 - 7c, 10 - 7d, 18g - 24
tigE1:     F23f, 13 - 12c, 1 - 3, 5 - 10b, 4 - 3f, 10c - 12b
tigF1:     1a-d, 6 - 1e, 7 - 10, 17 - 11, 18 - 20, 21 - 23e

Click here for the polytene chromosomes.

Pupal spur with about 5 or 6 spines.

Found: Ontario - Clarke Lake, Algonquin Provincial Park
            Quebec - Lake St. Joseph
            Minnesota - Spearhead Lake, Beltrami Co.; Turtle Lake, Becker Co.
            Wisconsin - Friebauer Lake, Bayfield Co.

            In thick mud at depths around 6 m in lakes.

Cytology mentioned in Martin et al. (1974) and Martin (1979), described by Kiknadze et al. (1993) as C. sp. Am1.    Redescribed and named by Butler & Kiknadze (in preparation).    Corresponds to Species 1 of Wülker.
It may be worthwhile checking the adults of this species against the type specimens of C. imperator Walley (1926).

[ Return to Index ]