Adult: Male Ð Dark species, thorax almost black, abdomen brown to dark brown, tergites uniformly colored. Sternite I with about 11 - 36 setae, sternite II with 2 - 6 lateral and about 70 medial setae. Ratio of 3rd-5th maxillary palpomeres (µm) 416 : 386 : 411. AR 5.8 - 7.3.
Superior volsella of the E-type, with thickening on the inner side. Inferior volsellae conical. Gonostylus broad, usually uniformly colored but sometimes with an obvious light area on external side. Anal point brownm narrowing from base to top.
Anterior basitarsus with beard, BR 5.25 - 6.78. Anterior LR 1.03 - 1.08. Sensilla chaetica on L2, 22 - 29.
Pupa: Thorax dark brown, tergites of abdomen with grey spots, congruent with shagreen areas. Laterosternites dark brown. Abdomen length about 9.9 - 11.6 mm. Wing sheath length about 2.58 - 3.02 mm.
Lateral spur of segment VIII almost black, usually with one spine, but sometimes with up to three, including a second tip.
Larva a large salinarius-type, length up to 20 mm. Gular region, frontoclypeal and other dorsal areas of head darkened; in some specimens there may be a pale spot on the posterior half of the frontoclypeus. Mentum of only specimen worn; c2 teeth well separated (type II), fourth laterals appear reduced, but in Russian specimens it appears only slightly reduced (Shobanov et al. 2002). AR 1.91 - 2.34. Pecten epipharyngis with 11 - 116 teeth. Mandible with 3rd inner tooth pale (type II), 12 - 15 striae on inner surface near the base.
Cytology
: 4 pairs of chromosomes with the thummi arm combination AB, CD, EF, G.
Arm G partly unpaired and relatively short (about 20 bands), with a virtually terminal
nucleolus and a Balbiani ring. Nucleolus median in arm D (group 9) and a small nucleolus in arm A in group 15, near the centromere. Wülker
(pers. comm.) notes relationship to C. neocorax.
traA1: 1a-2c, 10-12, 3i-d,13f-a, 4-9, 2d-3c, 14-19 ie. as holomelas
2
traA2: 1-2c, 10-12, 3ih, 8d-4, 13a-f, 3d-g, 8e-9, 2d-3c, 14-19
traB: with the typical position of groups 8-9 in the middle of the arm, flanked distally by dark band groups. The Ellesmere specimen has B1.1.
traC1: 1-3, 8-11c, 13-15, 4-6b, 12-11d, 6gh, 17a-16, 7-6c, 17b-22 i.e. as heteropilicornis
traD1: 1 - 3g, 11 - 18d, 9 - 10, 4 - 8, 18e-24; nucleolus in region 9.
traE1: 1a-c, 2e - 3e, 5 - 8g, 2d - 1d, 8h - 10b, 4 - 3f, 10c - 13 i.e. as albimaculatus
traF1: 1 - 9f, 11h - 17, 10d-a, 11g-a, 18 -23
Click here for polytene chromosome arms A, F and G or C-E.
Found: North West Territories - Skeleton Lake, Hazen, Ellesmere Island.
Alaska - west shore of Prudhoe Bay, EWD1, tundra pond east of West Doc pad.
Russia - Ice Sea coast: Lena Delta, Dikson, Archangelsk.
Cytology partially described by Wülker & Martin (2000). Full cytological and morphological description by Shobanov et al. (2002)