3. Chironomus analis Freeman, 1959.

Description of C. analis from Freeman 1959

Adult: From a single available specimen:
AR about 3.3.    Frontal tubercles about 260 microns.    About 26 clypeal setae.
At least 15 achrostichal setae; 18 dorsocentrals; 6 - 7 prealars; scutellars in two rows - anterior row about 9, posterior row about 13 setae.
Leg lengths (microns) and proportions as follows:

	Fe	Ti	Ta1	Ta2	Ta3
PI	1800	1760	2440	1400	1020
PII	1960	1880	1160	720	520
PIII	2280	2360	1600	1000	720
	Ta4	Ta5	LR	F/T	BR
PI	880	440	1.39	1.02	4.8
PII	360	240	0.62	1.04
PIII	480	280	0.68	0.97

Abdomen:    Segments 2-5 greenish with dark saddle spots in the basal region of the segment.    9 setae on 9th tergite.    Anal point broad and stout, superior volsella curved.

Pupa: From notes by Don Forsyth, the pupa has a dark exuvia, with dark muscle scars, and a very dark spur with multiple appressed spines.

Larval Morphology:  salinarius-type larva.    Head capsule coloration similar to that of C. zealandicus, but a smaller species, length about 15.3 -19.0 mm (female), 10.7 - 13.2 (male).     Anal tubules short and rounded or pointed, less than 3 times longer than wide.     Mentum (Fig. c) generally type III; centre tooth with C2 teeth well separated(i.e. type II).    Ventromental plate (Fig. d) with only about 30 - 36 striae.    Pecten epipharyngis (Fig. a) with about 8 - 14 teeth, some reduced.    Basal segment of antenna (Fig. b) not as long and narrow as in other NZ species such as C. zealanicus and , about 2.8 - 3.5 times as long as wide (similar to C. species 6).    Antennal proportions (microns) 132 : 32 : 7 : 15 : 7 ; AR 1.88 - 2.3.    Mandible (Fig. e) type II, with about 13 - 25 striae near base.

Cytology:  4 polytene chromosomes, pseudothummi-complex arm combination (BF,CD,AE, G).
Two nucleoli; a large one in arm F and a small one in arm G, often very difficult to distinguish from a BR.    Two BRs towards the other end of the arm.    Arm G usually only partially paired, sometimes more so than in other cells or specimens.    Arm A as sequence A4 in Australian species; anlE1 with sequence E1 as Australian species; anlF1 as F3 in Australian species.    Polymorphism at least in arms B, C and F.

anlA1:    1a-e, 11 - 10, 2c - 1f, 3e - 2d, 8 - 9, 3f-i, 12c-a, 4 - 7, 13 - 19                ie. as oppositus A4, forsythi
anlB1:    Puff (group 7) in proximal third of arm, possibly as forB3.
anlB2:    inversion of distal half of arm, seen as heterozygote at Lake Ngapouri.
anlC1:    as oppositus C4?
anlC2:    as oppositus C3?    seen at Lake Ngapouri.
anlD1:    1-2, 16 - 13, 8- 3e, 9, 3d-a, 10d - 12, 10a-c, 17 - 24
anlE1:    1 - 3e, 10b - 3f, 10c - 13                                                    ie. as oppositusE1, forsythi
anlF1: (approx.) 1 - 2a, 10 - 2c, 15c - 11a, 2b, 15d - 23                    ie. as oppositus F3, australis, forsythi, novae-zelandiae
anlF2:    1 - 2a, 10 - 9c, 2c - 9b, 15c - 11, 2b, 15d - 23
anlG1:    essentially as forsythi ?

Click here to see the polytene chromosomes

Known Localities

Some information on larval morphology and cytology given by Martin (1998).    Stark (1981) in his key to larvae, has C. analis as a thummi-type larva, but the basis for this conclusion is not known.

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