Redescribed by Hutton 1902.
C. species a.(Forsyth, 1978).
Hutton's description, aside from a detailed colour description, provides the information that length is 6 - 8 mm, wing length about 5.5 mm, and LR about 1.3.
Additional specimens:
AR about 3.6 - 4.0.
Wing length 4.6 - 5.5 mm; width 0.96 - 1.16 mm.; VR 0.95.
Cephalic tubercles present 10 - 35 micron.
Setae: 16 - 33 Clypeal; 16 - 20 Dorsocentrals; abt 6 prealars; 17 - 22 Scutellar, sometimes in two rows (posterior row larger) sometimes in multiple rows (anterior rows more numerous than posterior row).
LR 1.3 - 1.44; Anterior tarsi with a short dense beard (BR about 3 - 6) and sparse longer hairs (BR about 6 - 9).
Leg lengths (microns) and proportions:
|
|
Fe |
Ti |
Ta1 |
Ta2 |
Ta3 |
|
PI |
1600 |
1540 |
1980 |
1230 |
870 |
|
PII |
1710 |
1720 |
960 |
650 |
480 |
|
PIII |
2110 |
2200 |
1430 |
940 |
700 |
|
|
Ta4 |
Ta5 |
LR |
F/T |
BR |
|
PI |
700 |
350 |
1.29-1.44 |
1.03-1.08 |
6-9 |
|
PII |
320 |
230 |
0.55-0.59 |
0.96-1.08 |
|
|
PIII |
460 |
270 |
0.65-0.70 |
0.94-1.00 |
|
Abdominal tergites with a broad dark band covering most of the surface, leaving only a narrow pale band at the posterior edge.
About 8 - 15 setae at base of anal point.
Pupa Length about 10.2 - 12.5 mm (males), inner margin of wing case about 2.45 mm (males). Cephalic tubercle about 134 - 160 micron, with sub-apical seta about 78 - 88 micron. Muscle scars on abdominal tergites slightly darkened; about 65 - 100 recurved hooks on abdominal segment II, spurs with numerous spines (18 - 22) spreading out like a hand with the fingers spread. About 110 - 115 multiple ranked setae on each side of the anal lobe.
Larval Morphology: salinarius-type larva. Largest NZ species, length about 15.3 - 21.5 mm. Head capsule with frontoclypeus and most of the gular region very dark. Anal tubules short and often tending to a point. Most distinctive feature is the high number of striae (75 - 99) on the Ventromental plate. Mentum of type II (i.e 4th laterals reduced at least to the level of 5th laterals) with central trifid tooth of Type II, although sometimes the c2 teeth are less separated from the c1 tooth. Pecten epipharyngis with about 12 - 15 relatively sharp teeth, some reduced. Antennal segments in proportions (micron) 163 : 37 : 8 : 15 : 8 ; AR 2.0 - 2.3; basal segment of antenna relatively long and narrow, about 3.1 - 3.7 times as long as wide. Premandible with the usual two blades, inner blade about twice the width of the outer (2.1). Mandible with 3rd inner tooth darkened and often fully separate (i.e. Type III); about 17 - 23 striae on the outer surface near the base.
Cytology: 3 polytene chromosomes, modified pseudothummi-complex arm combination (BF,CD,GEA).
Three nucleoli, one in arm G, one proximal and another medial in arm F(Lentzios & Stocker,1979). The medial
nucleolus may not always be developed, since Lentzios & Stocker (1979) did not find it in their material. zeaA1 differing from oppositus A4 sequence by a simple inversion; arm E pattern not typical but zeaE1 appears to differ by a small inversion 5c-7 from oppositus E1. Arm F differs from that of novae-zealandiae, etc., by inversion of the region 7 to 11. Considerable polymorphism in arms A,
B,C, E, and G(?); sex determiner in arm C, probably near the centromere (Martin & Lee, 1984).
Lentzios et al. (1980) give some information on the number and location of C-bands.
zeaA1: 1a-e, 2g - 3, 1f - 2e, 10 - 11, 2f-d, 8 - 9, 3f-i, 12c-a, 4 - 7, 13 - 19
hypoth: 1a-e, 2g - 3, 4a, 12a-c, 3i-f, 9 - 8, 2d-f, 11 - 10, 2e - 1f, 3e-b, 4b - 7, 13 - 19
zeaA2: 1a-c, 10 - 11, 2f-d, 8 - 9, 3f-i, 12c-a, 4a, 3a - 2g, 1ed, 2e - 1f, 3e-b, 4b - 7, 13 - 19
zeaB1: Puff, with proximal dark bands (groups 7 and 8), in proximal third of arm.
zeaB2: Inversion of distal end of arm, distal to large puff.
zeaC1:
zeaC2: Inversion of about 2/3 of arm.
zeaD1: 1-2, 16-13, 18, 5-3e, 10a-c, 6-9, 3d-a, 10d-12, 19-24
zeaE1: (approx) 1?- 3e, 10b - 8, 5c - 7, 5b - 3f, 10c - 13
zeaE2: (approx) 1?-f, 4b - 5b, 7 - 5c, 8 - 10b, 3e - 2, 4a - 3f, 10c - 13
zeaE3: (approx) 1?- 3e, 10b - 8, 5c - 7, 5b - 3f, 10c-g, 12 -11, 13
zeaF1: 1 - 2a, 10 - 8, 11 - 15c, 2c - 7, 2b, 15d - 23 (In7 - 11 from oppositus F3)
zeaG1: fused to arm E.
zeaG2: possible inversion
Click here to see the polytene chromosomes
This appears to have been the "salinarius-type" studied by Robb (1966). It is referred to as C. zealandicus Type 2 by Lentzios & Stocker (1979) and Lentzios et al. (1980), who received their material from Lake Okaro.
Martin (1998) gives some information and drawings of immature stages.